{"id":31396,"date":"2023-06-01T00:13:46","date_gmt":"2023-05-31T16:13:46","guid":{"rendered":"https:\/newsletter.sinica.edu.tw\/?p=31396"},"modified":"2023-06-01T00:40:04","modified_gmt":"2023-05-31T16:40:04","slug":"%e3%80%90%e5%b0%88%e6%ac%84%e3%80%91%e6%a4%8d%e7%89%a9%e7%9a%84%e9%9d%a2%e5%ad%90-%e6%bc%ab%e8%ab%87%e6%a4%8d%e7%89%a9%e7%9a%84%e8%91%89%e8%a1%a8","status":"publish","type":"post","link":"https:\/newsletter.sinica.edu.tw\/31396\/","title":{"rendered":"\u3010\u5c08\u6b04\u3011\u690d\u7269\u7684\u9762\u5b50\u2014\u2014\u6f2b\u8ac7\u690d\u7269\u7684\u8449\u8868"},"content":{"rendered":"

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\u25b2\u5716\u4e8c \u6c23\u5b54\u8868\u578b\u56e0\u7269\u7a2e\u800c\u7570\uff1a\uff08A\uff09\u6c34\u7a3b\u4e2d\u7684\u555e\u9234\u5f62\u6c23\u5b54\uff08\u9ec3\u8272\uff09\uff1b\uff08B\uff09\u64ec\u5357\u82a5\u4e2d\u7684\u814e\u5f62\u6c23\u5b54\uff1b\uff08C\uff09Begonia hernandioides<\/em> \u4e2d\u7684\u7c07\u72c0\u6c23\u5b54\u3002<\/p><\/div>\n

\u70ba\u4e86\u4e86\u89e3\u7fa4\u805a\u578b\u6c23\u5b54\u662f\u5426\u80fd\u589e\u52a0\u4e86\u9678\u751f\u690d\u7269\u5728\u4e7e\u65f1\u74b0\u5883\u4e2d\u7684\u751f\u5b58\u512a\u52e2\uff0c\u6211\u5011\u5229\u7528\u5169\u7a2e\u4e9e\u6d32\u79cb\u6d77\u68e0\uff0cBegonia formosana<\/em>\uff08\u55ae\u4e00\u578b\u6c23\u5b54\uff09\u548c B. hernandioides<\/em>\uff08\u7fa4\u805a\u6c23\u5b54\u578b\uff09\u4f86\u63a2\u7d22\u690d\u7269\u7684\u6c34\u4efd\u4f7f\u7528\u80fd\u529b\u53ca\u7fa4\u805a\u578b\u6c23\u5b54\u7684\u767c\u80b2\u6a5f\u5236 7<\/sup>\u3002\u7814\u7a76\u767c\u73fe\u5728\u9ad8\u5149\u5f37\u5ea6\u4e0b\uff0cB. hernandioides<\/em> \u8868\u73fe\u51fa\u66f4\u9ad8\u7684\u6c34\u5206\u5229\u7528\u6548\u7387\uff0c\u6c23\u5b54\u958b\u5408\u7a0b\u5ea6\u8f03\u5c0f\uff0c\u958b\u5b54\u901f\u5ea6\u8f03\u5feb\u3002\u5206\u6790\u7fa4\u805a\u578b\u6c23\u5b54\u4e4b\u9593\u7684\u8ddd\u96e2\u767c\u73fe\uff0c\u7fa4\u805a\u578b\u6c23\u5b54\u53ef\u4ee5\u4fc3\u9032\u7d30\u80de\u9593\u7684\u4ea4\u4e92\u4f5c\u7528\uff0c\u4ee5\u5be6\u73fe\u6c23\u5b54\u904b\u52d5\u7684\u540c\u6b65\u6027\u3002\u53e6\u5916\uff0c\u7814\u7a76\u6291\u5236\u6c23\u5b54\u767c\u80b2\u76f8\u95dc\u57fa\u56e0 TMM\uff08too many mouths\uff09\uff0c\u4e5f\u767c\u73fe B. hernandioides<\/em> \u7684 TMM \u529f\u80fd\u6bd4\u8f03\u4f4e\u5f31\uff0c\u53ef\u80fd\u70ba\u7fa4\u805a\u578b\u6c23\u5b54\u767c\u751f\u7684\u539f\u56e0\u3002\u79cb\u6d77\u68e0\u6c23\u5b54\u805a\u96c6\u53ef\u80fd\u662f\u4e00\u7a2e\u751f\u5b58\u7b56\u7565\uff0c\u4ea6\u5373\u5efa\u7acb\u66f4\u5c0f\u66f4\u8fd1\u7684\u6c23\u5b54\u4ee5\u52a0\u901f\u5c0d\u5149\u7684\u53cd\u61c9\uff0c\u6e1b\u5c11\u6c34\u4efd\u6563\u5931\u3002<\/p>\n

\u5728\u690d\u7269\u9678\u751f\u5316\u7684\u904e\u7a0b\u4e2d\uff0c\u89d2\u8cea\u7684\u751f\u6210\u53ca\u6c23\u5b54\u7684\u7522\u751f\u7686\u70ba\u690d\u7269\u9632\u6b62\u53ca\u8abf\u7bc0\u6c34\u4efd\u6563\u5931\u7684\u91cd\u8981\u7279\u5fb5\u3002\u7136\u800c\u6b64\u4e8c\u8005\u5728\u767c\u80b2\u4e0a\u662f\u5426\u6709\u76f8\u95dc\u4e26\u4e0d\u6e05\u695a\u3002\u5728\u4e00\u4efd 1972 \u5e74\u7684\u7814\u7a76\u66fe\u7d93\u63d0\u53ca\u4e00\u7a2e\u5927\u9ea5\u7684\u7a81\u8b8a\u9ad4\u64c1\u6709\u8f03\u5c11\u7684\u89d2\u8cea\u5c64\u4e14\u6709\u6c23\u5b54\u7fa4\u805a\u7684\u73fe\u8c61\uff0c\u4f46\u6f5b\u5728\u7684\u767c\u80b2\u6a5f\u5236\u53ca\u6b64\u4e8c\u8005\u53c8\u662f\u5982\u4f55\u4e92\u76f8\u8abf\u63a7\uff0c\u90fd\u672a\u66fe\u88ab\u63a2\u8a0e\u904e\u3002\u6211\u5011\u5229\u7528\u5206\u5b50\u751f\u7269\u5b78\u7684\u7814\u7a76\u65b9\u5f0f\u627e\u51fa\u4e86\u8abf\u63a7\u8449\u8868\u76ae\u89d2\u8cea\u751f\u6210\u7684\u57fa\u56e0\uff0c\u4e26\u767c\u73fe\u5728\u8449\u5b50\u767c\u80b2\u7684\u65e9\u671f\uff0c\u6c23\u5b54\u7684\u539f\u59cb\u7d30\u80de\u5fc5\u9808\u6291\u5236\u89d2\u8cea\u7684\u5408\u6210\uff0c\u5373\u9019\u6642\u7684\u7d30\u80de\u5fc5\u9808\u5920\u67d4\u8edf\u4ee5\u7dad\u6301\u8868\u76ae\u5c64\u7684\u9069\u7576\u7d50\u69cb\u548c\u751f\u7269\u529b\u5b78\uff0c\u4ee5\u5f62\u6210\u9069\u7576\u7684\u6c23\u5b54\u6392\u5217 8<\/sup>\u3002\u9019\u8aaa\u660e\u4e86\u9664\u4e86\u751f\u7269\u5316\u5b78\u4fe1\u865f\u4e4b\u5916\uff0c\u8868\u76ae\u7684\u7269\u7406\u6027\u5f35\u529b\u4e5f\u6703\u5f71\u97ff\u7d44\u7e54\u7684\u751f\u9577\u8207\u767c\u80b2\uff0c\u800c\u62fc\u5716\u72c0\u8449\u8868\u76ae\u7d30\u80de\u7684\u5f62\u6210\u4e5f\u662f\u5176\u4e2d\u4e00\u500b\u4f8b\u5b50\u3002\u5229\u7528\u6a21\u64ec\u7684\u65b9\u5f0f\u63a8\u8ad6\u8868\u76ae\u7d30\u80de\u9577\u5927\u8207\u8449\u7247\u751f\u9577\u65b9\u5411\u7684\u76f8\u4e92\u59a5\u5354\u4e0b\uff0c\u7d30\u80de\u5169\u5169\u9472\u5d4c\u53ef\u5f62\u6210\u6700\u7a69\u5b9a\u7d50\u69cb\u7684\u7d50\u679c 9<\/sup>\uff1b\u6b64\u767c\u73fe\u53ef\u9a57\u8b49\u5169\u767e\u5e74\u524d\u690d\u7269\u5b78\u5bb6\u6240\u63d0\u51fa\u7684\u91cd\u8981\u767c\u80b2\u5047\u8aaa\uff1a\u8868\u76ae\u70ba\u6c7a\u5b9a\u7d44\u7e54\u5927\u5c0f\u7684\u91cd\u8981\u56e0\u7d20\uff08epidermal growth control theory, 1867\uff09 10<\/sup>\u3002<\/p>\n

\u690d\u7269\u4e0d\u6703\u79fb\u52d5\uff0c\u70ba\u4e86\u9069\u61c9\u5404\u7a2e\u74b0\u5883\uff0c\u5b83\u5011\u53ea\u80fd\u9760\u8457\u91cd\u5851\u81ea\u6211\u7684\u767c\u80b2\u800c\u5f97\u4ee5\u751f\u5b58\u8207\u7e41\u884d\u5f8c\u4ee3\u3002\u4f8b\u5982\uff0c\u8449\u8868\u76ae\u7d30\u80de\u5177\u6709\u9ad8\u5ea6\u53ef\u5851\u6027\uff0c\u5b83\u5011\u80fd\u5920\u91cd\u5851\u767c\u80b2\u904e\u7a0b\u4ee5\u7522\u751f\u4e0d\u540c\u7684\u8868\u76ae\u7d30\u80de\u7d44\u6210\u8207\u7d50\u69cb\u4f86\u56e0\u61c9\u74b0\u5883\u8b8a\u5316\uff0c\u56e0\u6b64\u89c0\u5bdf\u4e0d\u540c\u6642\u671f\u7684\u690d\u7269\u6a19\u672c\u6642\uff0c\u61c9\u53ef\u770b\u51fa\u690d\u7269\u8868\u76ae\u7d30\u80de\u56e0\u61c9\u7576\u4e0b\u5730\u7403\u74b0\u5883\u6240\u7522\u751f\u7684\u767c\u80b2\u8b8a\u5316\u3002\u4e00\u4efd 1987 \u5e74\u7684\u7814\u7a76\u6307\u51fa\uff0c\u5341\u516b\u4e16\u7d00\u5de5\u696d\u9769\u547d\u524d\u5927\u6c23\u4e8c\u6c27\u5316\u78b3\u542b\u91cf\u70ba 280 ppm \uff0c\u800c\u7576\u6642\u70ba 340 ppm\uff0c\u5169\u767e\u5e74\u9593\uff0c\u690d\u7269\u6c23\u5b54\u7684\u5bc6\u5ea6\uff08\u55ae\u4f4d\u9762\u7a4d\u4e0b\u7684\u6578\u91cf\uff09\u4e0b\u964d\u4e86 40% 11<\/sup>\uff0c\u7531\u6b64\u53ef\u898b\uff0c\u6c23\u5b54\u767c\u80b2\u662f\u5177\u6709\u53ef\u5851\u6027\u7684\u3002<\/p>\n

\u5728\u8449\u8868\u76ae\u7d30\u80de\u4e2d\uff0c\u6211\u5011\u628a\u6b64\u7a2e\u5177\u9078\u64c7\u589e\u751f\u6216\u5206\u5316\u6210\u7279\u5b9a\u529f\u80fd\u7684\u7d30\u80de\u7a31\u4e4b\u70ba\u5e79\u7d30\u80de\uff0c\u7136\u800c\u8449\u8868\u76ae\u5e79\u7d30\u80de\u7684\u5168\u80fd\u6027\uff08multipotency\uff09\u662f\u5982\u4f55\u7dad\u6301\u7684\uff1f\u5e79\u7d30\u80de\u53c8\u662f\u5982\u4f55\u6c7a\u5b9a\u8981\u7e7c\u7e8c\u5206\u88c2\u6216\u662f\u5206\u5316\uff1f\u4ecd\u5b58\u5728\u8a31\u591a\u672a\u77e5\u3002\u56e0\u6b64\u6211\u5011\u5229\u7528\u690d\u7269\u8449\u8868\u76ae\u7684\u5e79\u7d30\u80de\uff08SLGCs, stomatal lineage ground cells\uff09\u4f86\u63a2\u8a0e\u8449\u8868\u76ae\u5e79\u7d30\u80de\u7684\u5168\u80fd\u6027\uff0c\u7814\u7a76\u767c\u73fe\u5177\u5206\u88c2\u80fd\u529b\u7684\u8449\u8868\u76ae\u5e79\u7d30\u80de\u80fd\u85c9\u7531\u8abf\u63a7\u7d30\u80de\u9031\u671f\u6c7a\u5b9a\u5176\u589e\u751f\u6216\u662f\u5206\u5316 12<\/sup>\u3002\u7576\u4eca\u7684\u5927\u6c23\u4e8c\u6c27\u5316\u78b3\u542b\u91cf\u7d04\u70ba 411 ppm\uff0c\u52a0\u4e0a\u6975\u7aef\u6c23\u5019\u7684\u5f71\u97ff\uff0c\u5982\u4f55\u4f7f\u4f5c\u7269\u751f\u9577\u5f97\u66f4\u597d\u662f\u7576\u4ee3\u79d1\u5b78\u5bb6\u5171\u540c\u81f4\u529b\u7684\u76ee\u6a19\u3002\u5229\u7528\u5c0d\u65bc\u8449\u8868\u76ae\u5e79\u7d30\u80de\u7684\u4e86\u89e3\uff0c\u5c07\u53ef\u61c9\u7528\u65bc\u6539\u5584\u690d\u7269\u7684\u751f\u9577\u53ca\u751f\u7522\u529b\uff0c\u4ee5\u7a81\u7834\u6c23\u5019\u8b8a\u9077\u5c0d\u7ce7\u98df\u4f5c\u7269\u751f\u9577\u8207\u690d\u7269\u751f\u614b\u5e36\u4f86\u7684\u4e0d\u5229\u5f71\u97ff\u3002<\/p>\n

\n
    \n
  1. Chater, C. C. et al.<\/em> Origin and function of stomata in the moss Physcomitrella patens. Nature Plants<\/em> 2, 16179 (2016). https:\/doi.org:10.1038\/nplants.2016.179<\/a><\/li>\n
  2. Papanatsiou, M. et al.<\/em> Optogenetic manipulation of stomatal kinetics improves carbon assimilation, water use, and growth. Science<\/em> 363, 1456 (2019). https:\/doi.org:10.1126\/science.aaw0046<\/a><\/li>\n
  3. Caine, R. S. et al.<\/em> Rice with reduced stomatal density conserves water and has improved drought tolerance under future climate conditions. The New phytologist<\/em> (2018). https:\/doi.org:10.1111\/nph.15344<\/a><\/li>\n
  4. Raissig, M. T. et al.<\/em> Mobile MUTE specifies subsidiary cells to build physiologically improved grass stomata. Science<\/em> 355, 1215-1218 (2017). https:\/doi.org:10.1126\/science.aal3254<\/a><\/li>\n
  5. Lehmann, P. & Or, D. Effects of stomata clustering on leaf gas exchange. New Phytologist<\/em> 207, 1015-1025 (2015). https:\/doi.org:10.1111\/nph.13442<\/a><\/li>\n
  6. Gan, Y. et al.<\/em> Stomatal clustering, a new marker for environmental perception and adaptation in terrestrial plants. Botanical Studies<\/em> 51, 325-336 (2010).<\/li>\n
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  8. Yang, S. L., Tran, N., Tsai, M. Y. & Ho, C. K. Misregulation of MYB16 expression causes stomatal cluster formation by disrupting polarity during asymmetric cell divisions. The Plant cell<\/em> 34, 455-476 (2022). https:\/doi.org:10.1093\/plcell\/koab260<\/a><\/li>\n
  9. Sapala, A. et al.<\/em> Why plants make puzzle cells, and how their shape emerges. eLife<\/em> 7, e32794 (2018). https:\/doi.org:10.7554\/eLife.32794<\/a><\/li>\n
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  12. Ho, C.-M. K., Bringmann, M., Oshima, Y., Mitsuda, N. & Bergmann, D. C. Transcriptional profiling reveals signatures of latent developmental potential in Arabidopsis stomatal lineage ground cells. Proceedings of the National Academy of Sciences<\/em> 118, e2021682118 (2021). https:\/doi.org:10.1073\/pnas.2021682118<\/a><\/li>\n<\/ol>\n","protected":false},"excerpt":{"rendered":"

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